According to pollen analyses, such marshy plants as Alnus, Salix, Cryptomeria, Gramineae and Nymphaceae are abundant, Lepidobalanus, Castanea and Fraxinus are common, and Fagus cf.
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- 孢粉分析显示沼泽植物;如Alnus;Salix;Cryptomeria;Gramineae;Numphaceae丰富;Lepidobalanus;Castanea;Fraxinus常见;而Fagus cf.