Based on a molecular linkage map of the DH population,the two genes were mapped on chromosomes 1 and 11;

• 应用DH系已构建的分子图谱将这两对落粒性基因定位于第1染色体的G270-RG541和第11染色体的G320-RZ638之间,分别定名为sh-4和sh-5;