Fibrous thickenings are formed in the cells of the endothecium.In staminate flowers, the septum separating anther locules was cleaved and the stomium was degenerated to release the pollen grains.

• 在雄花，紧邻二个花药腔之隔膜细胞瓦解、唇细胞裂开，药隔组织分开，花粉粒即可释出；