The results of morphological anatomy show the lophis, anterior projections, posterior projections, bridges and rami of cingulum, arch of cingulum are variable distinctly to different species.

• 从形态学上来讲,生殖隔离取决于雌、雄外生殖器的构造。因而生殖器的形态构造在分类上具有重要的参考价值。