Sepals usually (rarely not) incrassate at base, veins usually obvious (rarely not); leaf blade variously shaped, margin and apex not incrassate, or margin scarious, apex spinose when blade aristate.

• 萼片通常(很少不)的增厚的在基部，脉通常明显(很少不)；叶片形状各异，不增厚的边缘和先端，或边缘干膜质，当叶片具芒时先端具刺。(13